Undergraduate Courses

Biochemistry and Biophysics (BIOL 101)

The study of life at the molecular level. Topics include the three-dimensional structures and function of large biological molecules, the human genome, and the design of antiviral drugs to treat HIV/AIDS.

The first of four modules in a yearlong foundational biology sequence; meets for the first half of the term.

Course Type: Undergraduate
Term: Spring , Term: Fall

Cell Bio & Membrane Physiology (BIOL 102)

The study of cell biology and membrane physiology. Topics include organization and functional properties of biological membranes, membrane physiology and signaling, rough endoplasmic reticulum and synthesis of membrane/secretory membrane proteins, endocytosis, the cytoskeleton, and cell division.

The second of four modules in a yearlong foundational biology sequence; meets for the second half of the term. Prerequisite: BIOL 101.

Course Type: Undergraduate
Term: Spring , Term: Fall

Genes and Development (BIOL 103)

Foundation principles for the study of genes, genetics, and developmental biology. How genes control development and disease; Mendel’s rules; examples of organ physiology.

The third of four modules in a yearlong foundational biology sequence; meets for the first half of the term. Prerequisites: BIOL 101, 102

Course Type: Undergraduate
Term: Spring , Term: Fall

Ecology & Evolutionary Biology (BIOL 104)

The study of evolutionary biology, animal behavior, and the history of life. Evolutionary transitions and natural selection. Adaptation at genic, chromosomal, cellular, organismal, and supra-organismal levels. Distributional and social consequences of particular suites of organismal adaptations.

The fourth of four modules in a yearlong foundational biology sequence; meets for the second half of the term. Prerequisites: BIOL 101, 102, and 103.

Course Type: Undergraduate
Term: Spring , Term: Fall

The Evolution of Beauty (E&EB 050)

Exploration of the fundamental mechanism of sexual selection and mate choice, and the patterns of display trait and mating preference coevolution. Additional topics include what happens when the freedom of mate choice is infringed or disrupted by sexual coercion or sexual violence; and the role of aesthetic evolution and sexual conflict in the evolution of human sexuality, pleasure, and sexual diversity. Enrollment limited to freshmen.

Preregistration required; see under Frist Year Seminar Program.

Professor: Richard Prum
Course Type: Undergraduate

Malaria Lyme & Vector Borne Disease (E&EB 106 )

Introduction to the biology of pathogen transmission from one organism to another by insects; special focus on malaria, dengue, and Lyme disease. Biology of the pathogens including modes of transmission, establishment of infection, and immune responses; the challenges associated with vector control, prevention, development of vaccines, and treatments.

Intended for non-science majors; preference to freshmen and sophomores. Prerequisite: high school biology.

Requires Permission of the Instructor

Course Type: Undergraduate
Term: Fall 2019
Day/Time: MW 1.00-2.15

Conservation Biology (E&EB 115)

An introduction to ecological and evolutionary principles underpinning efforts to conserve Earth’s biodiversity. Efforts to halt the rapid increase in disappearance of both plants and animals. Discussion of sociological and economic issues.

Professor: Linda Puth
Course Type: Undergraduate
Term: Fall 2019
Day/Time: MW 10.30-11.20, 1 HTBA

History of Life (E&EB 125)

Examination of fossil and geologic evidence pertaining to the origin, evolution, and history of life on Earth. Emphasis on major events in the history of life, on what the fossil record reveals about the evolutionary process, on the diversity of ancient and living organisms, and on the evolutionary impact of Earth’s changing environment.

Requires Permission of the Instructor

Course Type: Undergraduate
Term: Spring 2020
Day/Time: TTh 11.35-12.50

Plants and People (E&EB 145)

The interaction of plants and people throughout history explored from biological, historical, anthropological, and artistic perspectives. Basic botany, plants in the context of agriculture, trade and societal change, plants as inspiration, plants in the environment.  Includes field trips to the Marsh Botanical Garden greenhouse, Yale Peabody museum, Yale herbarium, Yale farm and one of the Yale art galleries

Professor: Linda Puth
Course Type: Undergraduate
Term: Spring 2020
Day/Time: MWF 10.30-11.20

Laboratory for Virus Discovery and Evolution (E&EB 175L)

An inquiry-based, hands-on introduction to sampling bacteriophages (bacteria-specific viruses) from natural environments. Emphasis on lab methods to characterize viruses via growth assays and genome sequencing, and to experimentally evolve viruses on bacteria. Readings and discussion on virus biodiversity, role of viruses in the environment, and virus applications to solve human problems.

Professor: Alita Burmeister
Course Type: Undergraduate
Term: Spring 2020
Day/Time: T 1:00 - 5:00

Intro Statistics: Life Sciences (E&EB 210)

Statistical and probabilistic analysis of biological problems, presented with a unified foundation in basic statistical theory. Problems are drawn from genetics, ecology, epidemiology, and bioinformatics.

Course Type: Undergraduate
Term: Fall 2018
Day/Time: TTh 1.00-2.15

General Ecology (E&EB 220)

The theory and practice of ecology, including the ecology of individuals, population dynamics and regulation, community structure, ecosystem function, and ecological interactions at broad spatial and temporal scales. Topics such as climate change, fisheries management, and infectious diseases are placed in an ecological context.

Prerequisite: MATH 112 or equivalent.

Professor: David Vasseur, Professor: Carla Staver
Course Type: Undergraduate
Term: Fall 2019
Day/Time: MWF 10.30-11.20

Laboratory for Evolution & Functional Traits (E&EB 223L)

Study of evolutionary novelties, their functional morphology, and their role in the diversity of life. Introduction to techniques used for studying the diversity of animal body plans. Evolutionary innovations that have allowed groups of organisms to increase their diversity.

https://www.youtube.com/watch?v=MbLVM7_Yu8s&feature=em-upload_owner#acti…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2020
Day/Time: T W Th 1:30 - 4:30

Evolutionary Biology (E&EB 225)

An overview of evolutionary biology as the discipline uniting all of the life sciences. Reading and discussion of scientific papers to explore the dynamic aspects of evolutionary biology. Principles of population genetics, paleontology, and systematics; application of evolutionary thinking in disciplines such as developmental biology, ecology, microbiology, molecular biology, and human medicine.

Professor: Paul Turner, Professor: Martha Muñoz
Course Type: Undergraduate
Term: Spring 2020
Day/Time: TTh 10.30-11.20, 1 HTBA

Ecology and Evolution of Infectious Disease (E&EB 228)

Overview of the ecology and evolution of pathogens (bacteria, viruses, protozoa) and their impact on host populations. Topics include theoretical concepts, ecological and evolutionary dynamics, molecular biology, and epidemiology of ancient and emerging diseases. 

Professor: Paul Turner
Course Type: Undergraduate

Field Ecology (E&EB 230)

A field-based introduction to ecological research, using experimental and descriptive approaches, comparative analysis, and modeling for field and small-group projects. Weekly field trips explore local lake, salt marsh, rocky intertidal, traprock ridge, and upland forest ecosystems. Includes one Saturday field trip and a three-day trip during the October recess.

Professor: Linda Puth
Course Type: Undergraduate

Evolution and Medicine (E&EB 235)

Introduction to the ways in which evolutionary science informs medical research and clinical practice. Diseases of civilization and their relation to humans’ evolutionary past; the evolution of human defense mechanisms; antibiotic resistance and virulence in pathogens; cancer as an evolutionary process. Students view course lectures on line; class time focuses on discussion of lecture topics and research papers.

Prerequisite: BIOL 101-104

Requires Permission of the Instructor

Professor: Stephen Stearns
Course Type: Undergraduate
Term: Fall 2019
Day/Time: TTh 11:35-12:50

Plant Diversity & Evolution (E&EB 246)

Introduction to the major plant groups and their evolutionary relationships, with an emphasis on the diversification and global importance of flowering plants.

Prerequisite: a general understanding of biology and evolution.

Professor: Erika Edwards
Course Type: Undergraduate
Term: Fall 2019
Day/Time: MW 1.00-2.15

Laboratory for Plant Diversity & Evolution (E&EB 247L)

Hands-on experience with the plant groups examined in the accompanying lectures. Local field trips.

Professor: Erika Edwards
Course Type: Undergraduate
Term: Fall 2019
Day/Time: T 1:00-4:00

Biology Of Terrestrial Arthropods (E&EB 250)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relationships, fossil record); physiology and functional morphology (water relations, thermoregulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Professor: Marta Wells
Course Type: Undergraduate
Term: Fall 2019
Day/Time: TTh 11.35-12.50

Laboratory for Biology Of Terrestrial Arthropods (E&EB 251L)

Comparative anatomy, dissections, identification, and classification of terrestrial arthropods; specimen collection; field trips.

Prerequisite: Concurrently with or after E&EB 250.

Professor: Marta Wells
Course Type: Undergraduate
Term: Fall 2019
Day/Time: W 1.30-4.30

Invertebrates I (E&EB 255)

A study of animal diversity, with a focus on the evolution of marine invertebrates. The course will draw extensively on the Invertebrate Zoology collections at the Peabody Museum. The lecture does not have to be taken concurrently with lab.

Professor: Casey Dunn
Course Type: Undergraduate
Term: Fall 2019
Day/Time: T Th 11.35 -12.50

Laboratory for Invertebrates I (E&EB 256L)

A study of animal diversity, with a focus on the evolution of marine invertebrates. The course will draw extensively on the Invertebrate Zoology collections at the Peabody Museum. The lecture and lab must be taken concurrently.

Professor: Casey Dunn
Course Type: Undergraduate
Term: Fall 2019
Day/Time: Th 1.30 - 4.30

Ichthyology (E&EB 264)

A survey of fish diversity, including jawless vertebrates, chimaeras and sharks, lungfishes, and ray-finned fishes. Topics include the evolutionary origin of vertebrates, the fossil record of fishes, evolutionary diversification of major extant fish lineages, biogeography, ecology, and reproductive strategies of fishes

Course Type: Undergraduate

Laboratory for Ichthyology (E&EB 265L)

Laboratory and field studies of fish diversity, form, function, behavior, and classification. The course primarily involves study of museum specimens and of living and fossil fishes.
 

Course Type: Undergraduate

Ornithology (E&EB 272)

An overview of avian biology and evolution, including the structure, function, behavior, and diversity of birds. The evolutionary origin of birds, avian phylogeny, anatomy, physiology, neurobiology, breeding systems, and biogeography.

Professor: Richard Prum
Course Type: Undergraduate
Term: Spring

Laboratory for Ornithology (E&EB 273L)

Laboratory and field studies of avian morphology, diversity, phylogeny, classification, identification, and behavior.

Requires Permission of the Instructor

Enrollment limited to 12.

Professor: Richard Prum
Course Type: Undergraduate

Biological Oceanography (E&EB 275)

Exploration of a range of coastal and pelagic ecosystems. Relationships between biological systems and the physical processes that control the movements of water and productivity of marine systems. Anthropogenic impacts on oceans, such as the effects of fishing and climate change. Includes three Friday field trips.

Requires Permission of the Instructor

Enrollment limited to 15.

Professor: Mary Beth Decker
Course Type: Undergraduate
Term: Spring 2020
Day/Time: T Th 11.35 -12.50

Mammalogy (E&EB 280)

The evolution and diversity of mammals, including primates. Origins, evolutionary history, systematics, morphology, biogeography, physiology, behavior, and ecology of major mammalian lineages. Accompanying laboratories focus on diagnostic morphological features of mammalian groups through examination of specimens from the Peabody Museum.

Professor: Eric Sargis
Course Type: Undergraduate

Comparative Developmental Anatomy of Vertebrates (E&EB 290)

A survey of the development, structure, and evolution of major vertebrate groups. Topics include the micro-anatomy of major organ systems, the developmental underpinnings of the vertebrate body plan, and the development, structure and evolution of the major organ systems such as the locomotory system, sensory organs, digestive tract, reproductive tract, and nervous system.

Professor: Günter Wagner
Course Type: Undergraduate
Term: Spring 2020
Day/Time: MWF 11.35-12.25

Laboratory for Comparative Anatomy of Vertebrates (E&EB 291L)

Microscopic examination of histological and embryological preparations. Dissection of selected vertebrate species including shark, bony fish, frog, lizard, and rat.

Requires Department Permission

To be taken with E&EB 290.

Professor: Zachary Lewis
Course Type: Undergraduate
Term: Spring 2020
Day/Time: Th 1.30-4.30, F 1:30-4:30

Comparative Physiology (E&EB 295)

Comparative focus on vertebrate animals and how individual organisms survive in their environments and how species deal with common problems (respiration, energy acquisition, reproduction) using similar, or sometimes very different, tools. Additional topics include specialized adaptations to extreme environments from high altitude to the deep seas and physiological mechanisms that facilitate survival at multiple levels: cells, tissues, organs, systems, and whole organisms.

Prerequisites Prerequisites: BIOL 101, 102 and CHEM 161, or permission of the instructor.

Professor: Zachary Lewis
Course Type: Undergraduate
Term: Fall 2019
Day/Time: MW 11.35-12.50

Primate Behavior and Ecology (E&EB 300)

Socioecology of primates compared with that of other mammals, emphasizing both general principles and unique primate characteristics. Topics include life-history strategies, feeding ecology, mating systems, and ecological influences on social organization.

Course Type: Undergraduate
Term: Fall 2019
Day/Time: T Th 9:00 - 10:15

Evolutionary Systems Biology (E&EB 310)

Introduction to the evolution of complex biological systems, including metabolism, gene regulatory networks and molecular structure and function. Course includes mathematical modeling and computer simulations of complex adaptive systems.

Prerequisites: MATH 115, BIOL 101, or permission of the instructor.

Professor: Alvaro Sanchez
Course Type: Undergraduate

Advanced Ecology (E&EB 320)

An advanced treatment of ecology, including species interactions, species coexistence theory, species-environment interactions, the maintenance and consequences of biological diversity, spatial ecology, food webs, and eco-evolutionary interactions.

Prerequisites: E&EB 220 and 225, or with permission of instructor.

Professor: David Post
Course Type: Undergraduate
Term: Spring 2020
Day/Time: TTh 1.00-2.15

Philosophy of Biology (E&EB 321)

This course is an introduction to the philosophy of biology, with application to specific current problems. It focuses on two major strands of thinking seeking answers to fundamental and complementary questions: “How do we know?” (epistemology) and “What things really exist in the world?” (ontology). These two themes have the most important impact on the practice of science, as they pertain to the nature of the scientific enterprise and how it works (epistemology and philosophy of science), as well as what scientists consider part of reality (=science related ontology: unicorns and phlogiston NO; atoms, electrons, YES; but what about species and genes? Do they have the same status as atoms?). In each of these fields of philosophy we outline the main positions and then discuss how they apply to past and current debates in biology, in particular, but not exclusively, evolutionary biology. Prerequisite: a semester of biology or a semester of philosophy. 1 Yale College course credit(s)

Professor: Casey Dunn, Professor: Günter Wagner
Course Type: Undergraduate
Term: Fall 2019
Day/Time: T 1.30-3.20

Limnology (E&EB 325)

Limnology, the study of the physical, chemical, and biological properties of inland waters, focuses on lakes where physical (light, temperature, and mixing) and chemical (dissolved elements and compounds) properties interact with the ecology and evolution of organisms. Topics include origins and morphology of inland waters; physical and chemical properties; diversity and interactions among the organisms found in lakes; historical perspectives; and understanding conservation and management in the context of global change. Frequent field trips to local freshwater ecosystems.

Prerequisites: E&EB 220 and E&EB 225, or with permission of instructor.

Professor: David Post
Course Type: Undergraduate
Term: Fall 2019
Day/Time: T 1.00 - 2.15, Th 1.00 - 5.00

Plant Structure and Function (E&EB 326)

This is an advanced botany course, preferably for students that have taken EEB 246 in addition to BIO 104; otherwise permission must be obtained from the instructor. A keen interest in plants is a must.

Professor: Erika Edwards
Course Type: Undergraduate

Culture and Human Evolution (E&EB 336)

Examination of the origins of human modernity in the light of evolutionary and archaeological evidence. Understanding, through a merger of evolutionary reasoning with humanistic theory, the impact of human culture on natural selection across the last 250,000 years. 1 Yale College course credit(s)

Course Type: Undergraduate
Term: Spring 2020
Day/Time: M 1.30-3.20

Primate Diversity and Evolution (E&EB 342)

The diversity and evolutionary history of living and extinct primates. Focus on major controversies in primate systematics and evolution, including the origins and relationships of several groups. Consideration of both morphological and molecular studies. Morphological diversity and adaptations explored through museum specimens and fossil casts.

Requires Permission of the Instructor

Recommended preparation: ANTH 116.

Professor: Eric Sargis
Course Type: Undergraduate

Life History Evolution (E&EB 380)

Life history evolution studies how the phenotypic traits directly involved in reproductive success are shaped by evolution to solve ecological problems. The intimate interplay between evolution and ecology.

Requires Permission of the Instructor

After E&EB 220 and 225, or with permission of instructor.

Professor: Stephen Stearns
Course Type: Undergraduate
Term: Spring 2020
Day/Time: TTh 11.35-12.50

Science of Complex Systems (E&EB 428)

Introduction to the quantitative analysis of systems with many degrees of freedom. Fundamental components in the science of complex systems, including how to simulate complex systems, how to analyze model behaviors, and how to validate models using observations. Topics include cellular automata, bifurcation theory, deterministic chaos, self-organized criticality, renormalization, and inverse theory.

Prerequisite: PHYS 301, MATH 247, or equivalent.

Course Type: Undergraduate
Term: Fall 2019
Day/Time: MWF 10.30-11.20

Human Osteology (E&EB 464)

A lecture and laboratory course focusing on the characteristics of the human skeleton and its use in studies of functional morphology, paleodemography, and paleopathology. Laboratories familiarize students with skeletal parts; lectures focus on the nature of bone tissue, its biomechanical modification, sexing, aging, and interpretation of lesions. 1 Yale College course credit(s)

Professor: Eric Sargis
Course Type: Undergraduate
Term: Spring 2020
Day/Time: T Th 1.00-2.15

Tutorial (E&EB 469)

Individual or small-group study for qualified students who wish to investigate an area of ecology or evolutionary biology not presently covered by regular courses. A student must be sponsored by a faculty member who sets requirements and meets weekly with the student. One or more written examinations and/or a term paper are required. To register, the student must submit a written plan of study approved by the faculty instructor to the director of undergraduate studies. Students are encouraged to apply during the term preceding the tutorial. Proposals must be submitted no later than the first day of the second week of the term in which the student enrolls in the tutorial. The final paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment. In special cases, with approval of the director of undergraduate studies, this course may be elected for more than one term, but only one term may be counted as an elective toward the requirements of the major. Normally, faculty sponsors must be members of the EEB department.

Requires Permission of the Instructor

http://eeb.yale.edu/sites/default/files/eeb_469_tutorial_form_rev_8.24.1…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2019, Term: Fall 2019
Day/Time: 1 HTBA

Senior Tutorial (E&EB 470)

Tutorial for seniors in the B.A. degree program who elect a term of independent study to complete the senior requirement. A student must be sponsored by a faculty member who sets requirements and meets weekly with the student. One or more written examinations and/or a term paper are required. To register, the student must submit a written plan of study approved by the faculty instructor to the director of undergraduate studies. Students are encouraged to apply during the term preceding the tutorial. Proposals must be submitted no later than the first day of the second week of the term in which the student enrolls in the tutorial. The final paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment. Normally, faculty sponsors must be members of the EEB department.

Requires Permission of the Instructor

Enrollment limited to seniors. Fulfills the senior requirement for the B.A. degree.

http://eeb.yale.edu/sites/default/files/eeb_470_senior_tutorial_form_rev…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2019, Term: Fall 2019
Day/Time: 1 HTBA

Research (E&EB 474)

One term of original research in an area relevant to ecology or evolutionary biology. This may involve, for example, laboratory work, fieldwork, or mathematical or computer modeling. Students may also work in areas related to environmental biology such as policy, economics, or ethics. The research project may not be a review of relevant literature but must be original. In all cases students must have a faculty sponsor who oversees the research and is responsible for the rigor of the project. Students are expected to spend ten hours per week on their research projects. Using the form available from the office of undergraduate studies or from the Classes server, students must submit a research proposal that has been approved by the faculty sponsor to the director of undergraduate studies, preferably during the term preceding the research. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The  final research paper is due in the hands of the of the director of of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

http://eeb.yale.edu/sites/default/files/eeb_474_research_form_rev_8.20.1…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2019, Term: Fall 2019
Day/Time: 1 HTBA

Senior Research (E&EB 475 / E&EB 476)

One term of original research in an area relevant to ecology or evolutionary biology. This may involve, for example, laboratory work, fieldwork, or mathematical or computer modeling. Students may also work in areas related to environmental biology such as policy, economics, or ethics. The research project may not be a review of relevant literature but must be original. In all cases students must have a faculty sponsor who oversees the research and is responsible for the rigor of the project. Students are expected to spend ten hours per week on their research projects. Using the form available from the office of undergraduate studies or from the Classes server, students must submit a research proposal that has been approved by the faculty sponsor to the director of undergraduate studies, preferably during the term preceding the research. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The final research paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

Enrollment limited to seniors. Fulfills the senior requirement for the B.S. degree.

http://eeb.yale.edu/sites/default/files/eeb_475_a_and_b_senior_research_…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2019, Term: Fall 2019
Day/Time: 1 HTBA

Intensive Senior Research (E&EB 495 / E&EB 496)

One term of intensive original research during the senior year under the sponsorship of a Yale faculty member. Similar to other research courses except that a more substantial portion of a student?s time and effort should be spent on the research project (a minimum average of twenty hours per week). A research proposal approved by the sponsoring faculty member must be submitted to the director of undergraduate studies; forms are available from the office of undergraduate studies. For research in the fall term, approval is encouraged during the spring term of the junior year. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The final research paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

One term of intensive research fulfills a portion of the senior requirement for the B.S. degree.

http://eeb.yale.edu/sites/default/files/eeb_495a_eeb_496b_intensive_sr_r…

Professor: Marta Wells
Course Type: Undergraduate
Term: Spring 2019, Term: Fall 2019
Day/Time: 1 HTBA