Graduate Courses

Undergraduate Courses | Graduate Courses | Online Course Information

In addition to the EEB courses listed on this site, A number of other departments offer courses that are appropriate for EEB grad students. These include MCDB, Geology and Geophysics, Anthropology, Statistics, and Yale Program in Organismal and Integrative Biology.

Advanced Topics in Ecology & Evolutionary Biology [Grad] (E&EB 500 / E&EB 501)

Topics to be announced. Graded Satisfactory/Unsatisfactory.

Course Type: Graduate
Term: Spring 2017, Term: Fall 2017
Day/Time: M 2:30 - 4:30

Advanced Topics in Ecology & Evolutionary Biology [Grad] (E&EB 501)

Topics to be announced. Graded Satisfactory/Unsatisfactory.

Course Type: Graduate
Term: Spring
Day/Time: M 2.30-4.30

Intro Statistics: Life Sciences (E&EB 510)

Statistical and probabilistic analysis of biological problems, presented with a unified foundation in basic statistical theory. Problems are drawn from genetics, ecology, epidemiology, and bioinformatics.

Professor: Walter Jetz
Course Type: Graduate
Term: Fall 2017
Day/Time: T Th 1:00 - 2:15

Conservation Biology [Grad] (E&EB 515)

An introduction to ecological and evolutionary principles underpinning efforts to conserve Earth’s biodiversity. Efforts to halt the rapid increase in disappearance of both plants and animals. Discussion of sociological and economic issues.

Course Type: Graduate
Term: Fall
Day/Time: MW 10.30-11.20, 1 HTBA

General Ecology [Grad] (E&EB 520)

The theory and practice of ecology, including the ecology of individuals, population dynamics and regulation, community structure, ecosystem function, and ecological interactions at broad spatial and temporal scales. Topics such as climate change, fisheries management, and infectious diseases are placed in an ecological context.

Course Type: Graduate
Term: Fall
Day/Time: MWF 10.30-11.20

Lab: Evolution & Functional Traits [Grad] (E&EB 523)

Study of evolutionary novelties, their functional morphology, and their role in the diversity of life. Introduction to techniques used for studying the diversity of animal body plans. Evolutionary innovations that have allowed groups of organisms to increase their diversity.

Course Type: Graduate
Term: Spring
Day/Time: TWTh 1.30-4.30

Evolutionary Biology [Grad] (E&EB 525)

An overview of evolutionary biology as the discipline uniting all of the life sciences. Evolution explains the origin of life and Earth’s biodiversity, and how organisms acquire adaptations that improve survival and reproduction. This course uses reading and discussion of scientific papers to emphasize that evolutionary biology is a dynamic science, involving active research to better understand the mysteries of life. We discuss principles of population genetics, paleontology, and systematics; application of evolutionary thinking in disciplines such as developmental biology, ecology, microbiology, molecular biology, and human medicine.

Course Type: Graduate
Term: Spring
Day/Time: TTh 10.30-11.20,1 HTBA

Ecology and Evolution of Infectious Disease [Grad] (E&EB 528)

Overview of the ecology and evolution of pathogens (bacteria, viruses, protozoa) and their impact on host populations. Topics include theoretical concepts, ecological and evolutionary dynamics, molecular biology, and epidemiology of ancient and emerging diseases. 

Course Type: Graduate
Term: Spring
Day/Time: TTh 10:30-11:20, 1HTBA

Field Ecology [Grad] (E&EB 530)

A field-based introduction to ecological research, using experimental and descriptive approaches, comparative analysis, and modeling for field and small-group projects. Weekly field trips explore local lake, salt marsh, rocky intertidal, traprock ridge, and upland forest ecosystems. Includes one Saturday field trip and a three-day trip during the October recess.

Course Type: Graduate
Term: Fall 2016
Day/Time: TTh 1:00-5:00

Evolution and Medicine [Grad] (E&EB 535)

Introduction to the ways in which evolutionary science informs medical research and clinical practice. Diseases of civilization and their relation to humans’ evolutionary past; the evolution of human defense mechanisms; antibiotic resistance and virulence in pathogens; cancer as an evolutionary process. Students view course lectures on line; class time focuses on discussion of lecture topics and research papers.

Course Type: Graduate
Term: Fall
Day/Time: Section 1: T 7.00-8.50p -or- Section 2: M 9.25-11.15

Responsible Conduct of Research [Grad] (E&EB 545)

Graded Satisfactory/Unsatisfactory.

Requires Permission of the Instructor

This 5-week discussion seminar considers issues related to the responsible conduct of research. Topics addressed include: research misconduct, plagiarism, data acquisition and management, mentoring and collaboration, authorship and peer review, the use of animals and humans in scientific research, sexual harassment, diversity, and balancing professional and personal life.

Course Type: Graduate
Term: Spring
Day/Time: M 2.30-4.30

Plant Diversity & Evolution [Grad] (E&EB 546)

Introduction to the major plant groups and their evolutionary relationships, with an emphasis on the diversification and global importance of flowering plants.

Course Type: Graduate
Term: Fall
Day/Time: MW 1.00-2.15

Lab: Plant Diversity & Evolution [Grad] (E&EB 547)

Hands-on experience with the plant groups examined in the accompanying lectures. Local field trips.

Course Type: Graduate
Term: Fall
Day/Time: T 1.00-5.00

Biology Of Terrestrial Arthropods [Grad] (E&EB 550)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relations, fossil record); physiology and functional morphology (water relations, thermo-regulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Course Type: Graduate
Term: Fall
Day/Time: TTh 11.35-12.50

Lab: Biology Of Terrestrial Arthropods [Grad] (E&EB 551)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relationships, fossil record); physiology and functional morphology (water relations, thermoregulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Course Type: Graduate
Term: Fall
Day/Time: W 1.30-4.30

Ichthyology [Grad] (E&EB 564)

A survey of fish diversity, including jawless vertebrates, chimaeras and sharks, lungfishes, and ray-finned fishes. Topics include the evolutionary origin of vertebrates, the fossil record of fishes, evolutionary diversification of major extant fish lineages, biogeography, ecology, and reproductive strategies of fishes.

Course Type: Graduate
Term: Fall
Day/Time: T Th 1:00-2:15

Laboratory for Ichthyology [Grad] (E&EB 565)

Laboratory and field studies of fish diversity, form, function, behavior, and classification. The course primarily involves study of museum specimens and of living and fossil fishes.
 

Course Type: Graduate
Term: Fall
Day/Time: W 1:30 - 4:30

Biological Oceanography [Grad] (E&EB 575)

Exploration of a range of coastal and pelagic ecosystems. Relationships between biological systems and the physical processes that control the movements of water and productivity of marine systems. Anthropogenic impacts on oceans, such as the effects of fishing and climate change. Includes three Friday field trips.

Course Type: Graduate
Term: Spring
Day/Time: TTh 11.35-12.50 1 HTBA

Clarifying connections: tradeoffs, pleiotropy, and other ideas (E&EB 604)

Stephen  Stearns

Professor: Stephen Stearns
Course Type: Graduate
Term: Fall
Day/Time: F 1:30-3:30

Evolutionary Functional Genomics, Cell Types and Homology [Grad] (E&EB 610)

Functional genomics has opened the opportunity to assess the activity state of all genes in the genomes in a largely scalable way. Many cell types, tissues and characters can readily be assessed across many species (if one has enough money), leading to a new field of evolutionary or comparative functional genomics. At the same time this new field of data analysis can be used to address many deep issues in organismic evolution, like the evolution of cell types, the homology among cell types etc.

In this seminar we will review the current state of published literature as it pertains to the evolutionary analysis of transcriptomes and epigenetic marks and their bearing on issues of cell and tissue evolution and homology.

Location: Yale Systems Biology Institute on Yale West Campus, Building ISTC (aka 850 West Campus Drive) room 101

Course Type: Graduate
Term: Fall
Day/Time: T 4:00 - 5:00

Evolutionary Ecological Genetics (E&EB 620)

Topics related to analyzing molecular genetic data to answer questions in evolution and ecology. Methods to detect selection in DNA sequences and other molecular data, and landscape genetics, overlaying genetic data on ecological maps from global imaging. Other topics will be determined by interests of participants.

Professor: Jeffrey Powell
Course Type: Graduate
Term: Fall 2017
Day/Time: Thursday, 4:00p.m. - 6:00p.m.

Advanced Ecology [Grad] (E&EB 624)

An advanced treatment of ecology, including species interactions, species coexistence theory, species-environment interactions, the maintenance and consequences of biological diversity, spatial ecology, food webs, and eco-evolutionary interactions.

Course Type: Graduate
Term: Spring 2017
Day/Time: T Th 1:00 - 2:15

Evolutionary Ecological Genetics (E&EB 624)

An introduction to conservation genetics for advanced undergraduates and graduate students. The importance of genetic diversity and the means for preserving it.

Professor: Jeffrey Powell
Course Type: Graduate
Term: Fall 2017
Day/Time: 3 HTBA

Biology of Insect Disease Vectors [Grad] (E&EB 650)

Insects transmit pathogens that cause many emerging and re-emerging human and agriculture-related diseases. Many of these diseases, which are referred to as neglected tropical diseases (NTDs), have a dramatically negative impact on human health in the developing world. Furthermore, they cause indirect devastation by significantly reducing agricultural productivity and nutrient availability, exacerbating poverty and deepening disparities. This course introduces students to the biological interactions that occur between major groups of important disease vectors and the pathogens they transmit. Lectures cover current research trends that relate to the ecology and physiology of insect vectors. Course content focuses on how these aspects of vector biology relate to the development and implementation of innovative and effective disease-control strategies.

Prerequisite: full year of college/university-level biology, or permission of the instructor(s).

Course Type: Graduate
Term: Spring
Day/Time: W 2.00- 2.50, Th 3.00-3.50

Ornithology [Grad] (E&EB 672)

An overview of avian biology and evolution, including the structure, function, behavior, and diversity of birds. The evolutionary origin of birds, avian phylogeny, anatomy, physiology, neurobiology, breeding systems, and biogeography.

Course Type: Graduate
Term: Spring
Day/Time: MWF 9.25-10.15

Laboratory for Ornithology [Grad] (E&EB 673)

Laboratory and field studies of avian morphology, diversity, phylogeny, classification, identification, and behavior.

Course Type: Graduate
Term: Spring
Day/Time: T 1.30-4.30

Life History Evolution [Grad] (E&EB 680)

Life history evolution studies how the phenotypic traits directly involved in reproductive success are shaped by evolution to solve ecological problems. Nowhere is the interplay between evolution and ecology more intimate.

Requires Permission of the Instructor

Course Type: Graduate
Term: Spring
Day/Time: TTh 11.35-12.50

Scientific Writing for Ecology and Evolutionary Biology (E&EB 725)

This course will provide guidance and practice for graduate students in grant and manuscript writing in the fields of ecology and evolutionary biology. Students will produce one grant application (NSF GRFP/DDIG or similar) and one manuscript for publication (on a topic of their choice, to contribute to their thesis or other ongoing work).

Professor: Carla Staver
Course Type: Graduate
Term: Fall 2017
Day/Time: Monday, 6:00p.m. - 8:00p.m.

Microbial Ecology and Evolution (E&EB 729)

This 1-credit graduate seminar course examines various topics in the ecology and evolution of microbes, with an emphasis on prokaryotes (Eubacteria, Archaea) and single-celled eukaryotes (yeasts, protists). The goal of this course is for the students to develop a quantitative understanding of microbial community ecology, focusing on the mechanisms of community assembly, microbial biogeography, and the role of evolution in structuring microbial communities and endowing them with function. Microbial communities have many unique properties and differ in fundamental ways from those formed by macroscopic organisms. The course will examine these unique properties through readings of the primary literature, including both classic papers and cutting-edge advances in the field. The course will be quantitative in nature, and will explicitly discuss how mathematical models can allow us to understand ecological and evolutionary processes in microbial communities.

Professor: Alvaro Sanchez
Course Type: Graduate
Term: Fall 2017
Day/Time: Wednesday, 1:00p.m. - 2:50p.m.

Long-term Temporal Dynamics of Ecological Systems [Grad] (E&EB 740)

Are ecological systems generally at (or near) equilibrium? Or are their transient dynamics so slow that we need to know how they behave far from equilibrium, too? This question is increasingly pressing in the face of ongoing global change – there remains substantial uncertainty about whether predictions based on ecological equilibria are relevant for predicting ecosystem responses to global change. For insight into this question, we will deal with temporal dynamics of ecosystems, integrating theoretical perspectives with both modern long-term ecological research and paleoecology. We will consider how theoretical concepts like characteristic time scales, lag and legacy effects, and cyclic behaviors apply to empirical work in real ecological systems.

Course Type: Graduate
Term: Spring
Day/Time: Th 10:00-12:00

Evolving Dynamical Systems [Grad] (E&EB 810)

An introduction to the ways evolving biological systems can be described, modeled, and analyzed by using a dynamical systems approach.  Concrete models will be explored with respect to field or laboratory observations.  Extensive use will be made of the software package Mathematica, but prior experience with the program is not required.

Course Type: Graduate
Term: Spring
Day/Time: T Th 9.00 - 10.15

Primate Diversity & Evolution [Grad] (E&EB 842)

The diversity and evolutionary history of living and extinct primates.  Focus on major controversies in primate systematics and evolution, including the origins and relationships of several groups.  Consideration of both morphological and molecular studies.  Morphological and diversity and adaptations explored through museum specimens and fossil casts.

Course Type: Graduate
Term: Spring
Day/Time: W 1:30-3:20

1st Year Intro Research & Rotation [Grad] (E&EB 900)

1st Year Intro Research & Rotation

Course Type: Graduate
Term: Spring , Term: Fall
Day/Time: 3 HTBA

2nd Year Research [Grad] (E&EB 950)

By arrangement with faculty.

Course Type: Graduate
Term: Spring , Term: Fall
Day/Time: 3 HTBA

Studies in Evolutionary Medicine I [Grad] (E&EB 960 )

The first term of a two-term course that begins in January. Students learn the major principles of evolutionary biology and apply them to issues in medical research and practice by presenting and discussing original papers from the current research literature. Such issues include lactose and alcohol tolerance; the hygiene hypothesis and autoimmune disease; human genetic variation in drug response and pathogen resistance; spontaneous abortions, immune genes, and mate choice; parental conflicts over reproductive investment mediated by genetic imprinting; life history trade-offs and the evolution of aging; the evolution of virulence and drug resistance in pathogens; the evolutionary genetics of humans and their pathogens; the ecology and evolution of disease; the evolutionary origin of diseases; and the emergence of new diseases. Students develop a research proposal based on one of their own questions in the spring term, spend the summer on a research project related to their research proposal, and write a paper based on the results of their research in the fall term. Credit and grades are awarded for each term. Only students who have engaged in summer research projects may enroll in the fall term.

Admission is by competitive application only.

http://eeb.yale.edu/sites/default/files/eeb_460b.pdf

Course Type: Graduate
Term: Spring
Day/Time: TTh 4.00-5.15

Studies in Evolutionary Medicine II [Grad] (E&EB 961)

Continuation of E&EB 960b. Students learn the major principles of evolutionary biology and apply them to issues in medical research and practice by presenting and discussing original papers from the current research literature. Such issues include lactose and alcohol tolerance; the hygiene hypothesis and autoimmune disease; human genetic variation in drug response and pathogen resistance; spontaneous abortions, immune genes, and mate choice; parental conflicts over reproductive investment mediated by genetic imprinting; life history trade-offs and the evolution of aging; the evolution of virulence and drug resistance in pathogens; the evolutionary genetics of humans and their pathogens; the ecology and evolution of disease; the evolutionary origin of diseases; and the emergence of new diseases. Students develop a research proposal based on one of their own questions in the spring term, spend the summer on a research project related to their research proposal, and write a paper based on the results of their research in the fall term. Credit and grades are awarded for each term. Only students who have engaged in summer research projects may enroll in the fall term.

Course Type: Graduate
Term: Fall
Day/Time: T 4.00-5.15 2 HTBA