Courses

This is a comprehensive listing of courses taught within EEB. Please consult the Online Course Information for a listing of courses offered this semester.

LINKS: Undergraduate Courses | Graduate Courses | Online Course Information

Biochemistry and Biophysics (BIOL 101)

The study of life at the molecular level. Topics include the three-dimensional structures and function of large biological molecules, the human genome, and the design of antiviral drugs to treat HIV/AIDS.

The first of four modules in a yearlong foundational biology sequence; meets for the first half of the term.

Course Type: Undergraduate
Term: Spring , Term: Fall

Cell Bio & Membrane Physiology (BIOL 102)

The study of cell biology and membrane physiology. Topics include organization and functional properties of biological membranes, membrane physiology and signaling, rough endoplasmic reticulum and synthesis of membrane/secretory membrane proteins, endocytosis, the cytoskeleton, and cell division.

The second of four modules in a yearlong foundational biology sequence; meets for the second half of the term. Prerequisite: BIOL 101.

Course Type: Undergraduate
Term: Spring , Term: Fall

Genes and Development (BIOL 103)

Foundation principles for the study of genes, genetics, and developmental biology. How genes control development and disease; Mendel’s rules; examples of organ physiology.

The third of four modules in a yearlong foundational biology sequence; meets for the first half of the term. Prerequisites: BIOL 101, 102

Course Type: Undergraduate
Term: Spring , Term: Fall

Ecology & Evolutionary Biology (BIOL 104)

The study of evolutionary biology, animal behavior, and the history of life. Evolutionary transitions and natural selection. Adaptation at genic, chromosomal, cellular, organismal, and supra-organismal levels. Distributional and social consequences of particular suites of organismal adaptations.

The fourth of four modules in a yearlong foundational biology sequence; meets for the second half of the term. Prerequisites: BIOL 101, 102, and 103.

Course Type: Undergraduate
Term: Spring , Term: Fall

Malaria Lyme & Vector Borne Disease (E&EB 106 )

Introduction to the biology of pathogen transmission from one organism to another by insects; special focus on malaria, dengue, and Lyme disease. Biology of the pathogens including modes of transmission, establishment of infection, and immune responses; the challenges associated with vector control, prevention, development of vaccines, and treatments.

Intended for non-science majors; preference to freshmen and sophomores. Prerequisite: high school biology.

Requires Permission of the Instructor

Course Type: Undergraduate
Term: Fall
Day/Time: MW 1.00-2.15

Conservation Biology (E&EB 115)

An introduction to ecological and evolutionary principles underpinning efforts to conserve Earth’s biodiversity. Efforts to halt the rapid increase in disappearance of both plants and animals. Discussion of sociological and economic issues.

Course Type: Undergraduate
Term: Fall
Day/Time: MW 10.30-11.20, 1 HTBA

History of Life (E&EB 125)

Examination of fossil and geologic evidence pertaining to the origin, evolution, and history of life on Earth. Emphasis on major events in the history of life, on what the fossil record reveals about the evolutionary process, on the diversity of ancient and living organisms, and on the evolutionary impact of Earth’s changing environment.

Requires Permission of the Instructor

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 11.35-12.50

Plants and People (E&EB 145)

The interaction of plants and people throughout history explored from biological, historical, anthropological, and artistic perspectives. Basic botany, plants in the context of agriculture, trade and societal change, plants as inspiration, plants in the environment.  Includes field trips to the Marsh Botanical Garden greenhouse, Yale Peabody museum, Yale herbarium, Yale farm and one of the Yale art galleries

Course Type: Undergraduate
Term: Spring
Day/Time: MWF 10.30-11.20

Virus Discovery and Evolution (E&EB 175)

An inquiry-based, hands-on introduction to sampling bacteriophages (bacteria-specific viruses) from natural environments. Emphasis on lab methods to characterize viruses via growth assays and genome sequencing, and to experimentally evolve viruses on bacteria. Readings and discussion on virus biodiversity, role of viruses in the environment, and virus applications to solve human problems.

Course Type: Undergraduate
Term: Fall
Day/Time: TBD

Intro Statistics: Life Sciences (E&EB 210)

Statistical and probabilistic analysis of biological problems, presented with a unified foundation in basic statistical theory. Problems are drawn from genetics, ecology, epidemiology, and bioinformatics.

Course Type: Undergraduate
Term: Fall
Day/Time: TTh 1.00-2.15

General Ecology (E&EB 220)

The theory and practice of ecology, including the ecology of individuals, population dynamics and regulation, community structure, ecosystem function, and ecological interactions at broad spatial and temporal scales. Topics such as climate change, fisheries management, and infectious diseases are placed in an ecological context.

Prerequisite: MATH 112 or equivalent.

Course Type: Undergraduate
Term: Fall
Day/Time: MWF 10.30-11.20

Evolution & Functional Traits Lab (E&EB 223L)

Study of evolutionary novelties, their functional morphology, and their role in the diversity of life. Introduction to techniques used for studying the diversity of animal body plans. Evolutionary innovations that have allowed groups of organisms to increase their diversity.

https://www.youtube.com/watch?v=MbLVM7_Yu8s&feature=em-upload_owner#acti…

Course Type: Undergraduate
Term: Spring
Day/Time: 3 HTBA

Evolutionary Biology (E&EB 225)

An overview of evolutionary biology as the discipline uniting all of the life sciences. Reading and discussion of scientific papers to explore the dynamic aspects of evolutionary biology. Principles of population genetics, paleontology, and systematics; application of evolutionary thinking in disciplines such as developmental biology, ecology, microbiology, molecular biology, and human medicine.

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 10.30-11.20, 1 HTBA

Ecology and Evolution of Infectious Disease (E&EB 228)

Overview of the ecology and evolution of pathogens (bacteria, viruses, protozoa) and their impact on host populations. Topics include theoretical concepts, ecological and evolutionary dynamics, molecular biology, and epidemiology of ancient and emerging diseases. 

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 10:30-11:20, 1HTBA

Field Ecology (E&EB 230)

A field-based introduction to ecological research, using experimental and descriptive approaches, comparative analysis, and modeling for field and small-group projects. Weekly field trips explore local lake, salt marsh, rocky intertidal, traprock ridge, and upland forest ecosystems. Includes one Saturday field trip and a three-day trip during the October recess.

Course Type: Undergraduate
2016
Term: Fall 2016
Day/Time: TTh 1:00-5:00

Evolution and Medicine (E&EB 235)

Introduction to the ways in which evolutionary science informs medical research and clinical practice. Diseases of civilization and their relation to humans’ evolutionary past; the evolution of human defense mechanisms; antibiotic resistance and virulence in pathogens; cancer as an evolutionary process. Students view course lectures on line; class time focuses on discussion of lecture topics and research papers.

Prerequisite: BIOL 101-104

Requires Permission of the Instructor

Course Type: Undergraduate
Term: Fall
Day/Time: Section 1: T 7.00-8.50p -or- Section 2: M 9.25-11.15

Plant Diversity & Evolution (E&EB 246)

Introduction to the major plant groups and their evolutionary relationships, with an emphasis on the diversification and global importance of flowering plants.

To be taken concurrently with E&EB 247L.

Prerequisite: a general understanding of biology and evolution.

Course Type: Undergraduate
Term: Fall
Day/Time: MW 1.00-2.15

Lab: Plant Diversity & Evolution (E&EB 247L)

Hands-on experience with the plant groups examined in the accompanying lectures. Local field trips.

To be taken concurrently with E&EB 246.

Course Type: Undergraduate
Term: Fall
Day/Time: T 1.00-5.00

Biology Of Terrestrial Arthropods (E&EB 250)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relationships, fossil record); physiology and functional morphology (water relations, thermoregulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Course Type: Undergraduate
Term: Fall
Day/Time: TTh 11.35-12.50

Lab: Biology Of Terrestrial Arthropods (E&EB 251L)

Comparative anatomy, dissections, identification, and classification of terrestrial arthropods; specimen collection; field trips.

Prerequisite: Concurrently with or after E&EB 250.

Course Type: Undergraduate
Term: Fall
Day/Time: W 1.30-4.30

Ichthyology (E&EB 264)

A survey of fish diversity, including jawless vertebrates, chimaeras and sharks, lungfishes, and ray-finned fishes. Topics include the evolutionary origin of vertebrates, the fossil record of fishes, evolutionary diversification of major extant fish lineages, biogeography, ecology, and reproductive strategies of fishes

Course Type: Undergraduate
Term: Fall
Day/Time: MW 1:00-2:15

Laboratory for Ichthyology (E&EB 265L)

Laboratory and field studies of fish diversity, form, function, behavior, and classification. The course primarily involves study of museum specimens and of living and fossil fishes.
 

Course Type: Undergraduate
Term: Fall
Day/Time: T 1:00-4:00

Ornithology (E&EB 272)

An overview of avian biology and evolution, including the structure, function, behavior, and diversity of birds. The evolutionary origin of birds, avian phylogeny, anatomy, physiology, neurobiology, breeding systems, and biogeography.

Course Type: Undergraduate
Term: Spring
Day/Time: MWF 9.25-10.15

Laboratory for Ornithology (E&EB 273L)

Laboratory and field studies of avian morphology, diversity, phylogeny, classification, identification, and behavior.

Requires Permission of the Instructor

Enrollment limited to 12.

Course Type: Undergraduate
Term: Spring
Day/Time: T 1.30-4.30

Biological Oceanography (E&EB 275)

Exploration of a range of coastal and pelagic ecosystems. Relationships between biological systems and the physical processes that control the movements of water and productivity of marine systems. Anthropogenic impacts on oceans, such as the effects of fishing and climate change. Includes three Friday field trips.

Requires Permission of the Instructor

Enrollment limited to 15.

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 11.35-12.50, 1 HTBA

Mammalogy (E&EB 280)

The evolution and diversity of mammals, including primates. Origins, evolutionary history, systematics, morphology, biogeography, physiology, behavior, and ecology of major mammalian lineages. Accompanying laboratories focus on diagnostic morphological features of mammalian groups through examination of specimens from the Peabody Museum.

Course Type: Undergraduate
Term: Fall
Day/Time: TTh 2:30-3:45

Comparative Developmental Anatomy of Vertebrates (E&EB 290)

A survey of the development, structure, and evolution of major vertebrate groups. Topics include the micro-anatomy of major organ systems, the developmental underpinnings of the vertebrate body plan, and the development, structure and evolution of the major organ systems such as the locomotory system, sensory organs, digestive tract, reproductive tract, and nervous system.

Course Type: Undergraduate
Term: Spring
Day/Time: MWF 11.35-12.25

Lab: Comparative Anatomy of Vertebrates (E&EB 291L)

Microscopic examination of histological and embryological preparations. Dissection of selected vertebrate species including shark, bony fish, frog, lizard, and rat.

Requires Department Permission

To be taken with E&EB 290.

Course Type: Undergraduate
Term: Spring
Day/Time: Section 1: Th 1.30-4.30 -or- Section 2: F 1.30-4.30

Advanced Ecology (E&EB 320)

An advanced treatment of ecology, including species interactions, species coexistence theory, species-environment interactions, the maintenance and consequences of biological diversity, spatial ecology, food webs, and eco-evolutionary interactions.

Prerequisites: E&EB 220 and 225, or with permission of instructor.

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 1.00-2.15

Primate Diversity and Evolution (E&EB 342)

The diversity and evolutionary history of living and extinct primates. Focus on major controversies in primate systematics and evolution, including the origins and relationships of several groups. Consideration of both morphological and molecular studies. Morphological diversity and adaptations explored through museum specimens and fossil casts.

Requires Permission of the Instructor

Recommended preparation: ANTH 116.

Course Type: Undergraduate
Term: Spring
Day/Time: 1 HTBA

Life History Evolution (E&EB 380)

Life history evolution studies how the phenotypic traits directly involved in reproductive success are shaped by evolution to solve ecological problems. The intimate interplay between evolution and ecology.

Requires Permission of the Instructor

After E&EB 220 and 225, or with permission of instructor.

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 11.35-12.50

Science of Complex Systems (E&EB 428)

Introduction to the quantitative analysis of systems with many degrees of freedom. Fundamental components in the science of complex systems, including how to simulate complex systems, how to analyze model behaviors, and how to validate models using observations. Topics include cellular automata, bifurcation theory, deterministic chaos, self-organized criticality, renormalization, and inverse theory.

Prerequisite: PHYS 301, MATH 247, or equivalent.

Course Type: Undergraduate
Term: Fall
Day/Time: MW 9.00-10.15

Studies in Evolutionary Medicine I (E&EB 460)

Principles of evolutionary biology applied to issues in medical research and practice. Lactose and alcohol tolerance; the “hygiene hypothesis”; genetic variation in drug response and pathogen resistance; spontaneous abortions, immune genes, and mate choice; the evolution of aging; the ecology and evolution of disease; the emergence of new diseases. Students develop proposals for research to be conducted during the summer.

Requires Permission of the Instructor

Admission by competitive application
http://eeb.yale.edu/sites/default/files/eeb_460b.pdf

Course Type: Undergraduate
Term: Spring
Day/Time: TTh 4.00-5.15

Studies in Evolutionary Medicine II (E&EB 461)

Continuation of E&EB 460.

Prerequisite: E&EB 460 or permission of instructor.

Course Type: Undergraduate
Term: Fall
Day/Time: TTh 4.00-5.15

Underclassman Tutorial (E&EB 469)

Individual or small-group study for qualified students who wish to investigate an area of ecology or evolutionary biology not presently covered by regular courses. A student must be sponsored by a faculty member who sets requirements and meets weekly with the student. One or more written examinations and/or a term paper are required. To register, the student must submit a written plan of study approved by the faculty instructor to the director of undergraduate studies. Students are encouraged to apply during the term preceding the tutorial. Proposals must be submitted no later than the first day of the second week of the term in which the student enrolls in the tutorial. The final paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment. In special cases, with approval of the director of undergraduate studies, this course may be elected for more than one term, but only one term may be counted as an elective toward the requirements of the major. Normally, faculty sponsors must be members of the EEB department.

Requires Permission of the Instructor

http://eeb.yale.edu/sites/default/files/eeb_469_tutorial_form_rev_8.24.1…

Course Type: Undergraduate
Term: Spring , Term: Fall
Day/Time: 1 HTBA

Senior Tutorial (E&EB 470)

Tutorial for seniors in the B.A. degree program who elect a term of independent study to complete the senior requirement. A student must be sponsored by a faculty member who sets requirements and meets weekly with the student. One or more written examinations and/or a term paper are required. To register, the student must submit a written plan of study approved by the faculty instructor to the director of undergraduate studies. Students are encouraged to apply during the term preceding the tutorial. Proposals must be submitted no later than the first day of the second week of the term in which the student enrolls in the tutorial. The final paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment. Normally, faculty sponsors must be members of the EEB department.

Requires Permission of the Instructor

Enrollment limited to seniors. Fulfills the senior requirement for the B.A. degree.

http://eeb.yale.edu/sites/default/files/eeb_470_senior_tutorial_form_rev…

Course Type: Undergraduate
Term: Spring , Term: Fall
Day/Time: 1 HTBA

Underclassman Research (E&EB 474)

One term of original research in an area relevant to ecology or evolutionary biology. This may involve, for example, laboratory work, fieldwork, or mathematical or computer modeling. Students may also work in areas related to environmental biology such as policy, economics, or ethics. The research project may not be a review of relevant literature but must be original. In all cases students must have a faculty sponsor who oversees the research and is responsible for the rigor of the project. Students are expected to spend ten hours per week on their research projects. Using the form available from the office of undergraduate studies or from the Classes server, students must submit a research proposal that has been approved by the faculty sponsor to the director of undergraduate studies, preferably during the term preceding the research. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The  final research paper is due in the hands of the of the director of of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

http://eeb.yale.edu/sites/default/files/eeb_474_research_form_rev_8.20.1…

Course Type: Undergraduate
Term: Spring , Term: Fall
Day/Time: 1 HTBA

Senior Research (E&EB 475)

One term of original research in an area relevant to ecology or evolutionary biology. This may involve, for example, laboratory work, fieldwork, or mathematical or computer modeling. Students may also work in areas related to environmental biology such as policy, economics, or ethics. The research project may not be a review of relevant literature but must be original. In all cases students must have a faculty sponsor who oversees the research and is responsible for the rigor of the project. Students are expected to spend ten hours per week on their research projects. Using the form available from the office of undergraduate studies or from the Classes server, students must submit a research proposal that has been approved by the faculty sponsor to the director of undergraduate studies, preferably during the term preceding the research. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The final research paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

Enrollment limited to seniors. Fulfills the senior requirement for the B.S. degree.

http://eeb.yale.edu/sites/default/files/eeb_475_a_and_b_senior_research_…

Course Type: Undergraduate
Term: Spring , Term: Fall
Day/Time: 1 HTBA

Intensive Senior Research (E&EB 495 / E&EB 496)

One term of intensive original research during the senior year under the sponsorship of a Yale faculty member. Similar to other research courses except that a more substantial portion of a student?s time and effort should be spent on the research project (a minimum average of twenty hours per week). A research proposal approved by the sponsoring faculty member must be submitted to the director of undergraduate studies; forms are available from the office of undergraduate studies. For research in the fall term, approval is encouraged during the spring term of the junior year. Proposals are due no later than the first day of the second week of the term in which the student enrolls in the course. The final research paper is due in the hands of the director of undergraduate studies by the last day of reading period in the term of enrollment.

Requires Permission of the Instructor

One term of intensive research fulfills a portion of the senior requirement for the B.S. degree.

http://eeb.yale.edu/sites/default/files/eeb_495a_eeb_496b_intensive_sr_r…

Course Type: Undergraduate
Term: Spring , Term: Fall
Day/Time: 1 HTBA

Advanced Topics in Ecology & Evolutionary Biology [Grad] (E&EB 500)

Topics to be announced. Graded Satisfactory/Unsatisfactory.

Course Type: Graduate
Term: Fall
Day/Time: M 2:00 - 4:00

Topics to be announced. Graded Satisfactory/Unsatisfactory.

Course Type: Graduate
Term: Spring
Day/Time: M 2.30-4.30

Conservation Biology [Grad] (E&EB 515)

An introduction to ecological and evolutionary principles underpinning efforts to conserve Earth’s biodiversity. Efforts to halt the rapid increase in disappearance of both plants and animals. Discussion of sociological and economic issues.

Course Type: Graduate
Term: Fall
Day/Time: MW 10.30-11.20, 1 HTBA

General Ecology [Grad] (E&EB 520)

The theory and practice of ecology, including the ecology of individuals, population dynamics and regulation, community structure, ecosystem function, and ecological interactions at broad spatial and temporal scales. Topics such as climate change, fisheries management, and infectious diseases are placed in an ecological context.

Course Type: Graduate
Term: Fall
Day/Time: MWF 10.30-11.20

Lab: Evolution & Functional Traits [Grad] (E&EB 523)

Study of evolutionary novelties, their functional morphology, and their role in the diversity of life. Introduction to techniques used for studying the diversity of animal body plans. Evolutionary innovations that have allowed groups of organisms to increase their diversity.

Course Type: Graduate
Term: Spring
Day/Time: TWTh 1.30-4.30

Evolutionary Biology [Grad] (E&EB 525)

An overview of evolutionary biology as the discipline uniting all of the life sciences. Evolution explains the origin of life and Earth’s biodiversity, and how organisms acquire adaptations that improve survival and reproduction. This course uses reading and discussion of scientific papers to emphasize that evolutionary biology is a dynamic science, involving active research to better understand the mysteries of life. We discuss principles of population genetics, paleontology, and systematics; application of evolutionary thinking in disciplines such as developmental biology, ecology, microbiology, molecular biology, and human medicine.

Course Type: Graduate
Term: Spring
Day/Time: TTh 10.30-11.20,1 HTBA

Ecology and Evolution of Infectious Disease [Grad] (E&EB 528)

Overview of the ecology and evolution of pathogens (bacteria, viruses, protozoa) and their impact on host populations. Topics include theoretical concepts, ecological and evolutionary dynamics, molecular biology, and epidemiology of ancient and emerging diseases. 

Course Type: Graduate
Term: Spring
Day/Time: TTh 10:30-11:20, 1HTBA

Field Ecology [Grad] (E&EB 530)

A field-based introduction to ecological research, using experimental and descriptive approaches, comparative analysis, and modeling for field and small-group projects. Weekly field trips explore local lake, salt marsh, rocky intertidal, traprock ridge, and upland forest ecosystems. Includes one Saturday field trip and a three-day trip during the October recess.

Course Type: Graduate
2016
Term: Fall 2016
Day/Time: TTh 1:00-5:00

Evolution and Medicine [Grad] (E&EB 535)

Introduction to the ways in which evolutionary science informs medical research and clinical practice. Diseases of civilization and their relation to humans’ evolutionary past; the evolution of human defense mechanisms; antibiotic resistance and virulence in pathogens; cancer as an evolutionary process. Students view course lectures on line; class time focuses on discussion of lecture topics and research papers.

Course Type: Graduate
Term: Fall
Day/Time: Section 1: T 7.00-8.50p -or- Section 2: M 9.25-11.15

Responsible Conduct of Research [Grad] (E&EB 545)

Graded Satisfactory/Unsatisfactory.

Requires Permission of the Instructor

This 5-week discussion seminar considers issues related to the responsible conduct of research. Topics addressed include: research misconduct, plagiarism, data acquisition and management, mentoring and collaboration, authorship and peer review, the use of animals and humans in scientific research, sexual harassment, diversity, and balancing professional and personal life.

Course Type: Graduate
Term: Spring
Day/Time: M 2.30-4.30

Plant Diversity & Evolution [Grad] (E&EB 546)

Introduction to the major plant groups and their evolutionary relationships, with an emphasis on the diversification and global importance of flowering plants.

Course Type: Graduate
Term: Fall
Day/Time: MW 1.00-2.15

Lab: Plant Diversity & Evolution [Grad] (E&EB 547)

Hands-on experience with the plant groups examined in the accompanying lectures. Local field trips.

Course Type: Graduate
Term: Fall
Day/Time: T 1.00-5.00

Biology Of Terrestrial Arthropods [Grad] (E&EB 550)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relations, fossil record); physiology and functional morphology (water relations, thermo-regulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Course Type: Graduate
Term: Fall
Day/Time: TTh 11.35-12.50

Lab: Biology Of Terrestrial Arthropods [Grad] (E&EB 551)

Evolutionary history and diversity of terrestrial arthropods (body plan, phylogenetic relationships, fossil record); physiology and functional morphology (water relations, thermoregulation, energetics of flying and singing); reproduction (biology of reproduction, life cycles, metamorphosis, parental care); behavior (migration, communication, mating systems, evolution of sociality); ecology (parasitism, mutualism, predator-prey interactions, competition, plant-insect interactions).

Course Type: Graduate
Term: Fall
Day/Time: W 1.30-4.30

Ichthyology [Grad] (E&EB 564)

A survey of fish diversity, including jawless vertebrates, chimaeras and sharks, lungfishes, and ray-finned fishes. Topics include the evolutionary origin of vertebrates, the fossil record of fishes, evolutionary diversification of major extant fish lineages, biogeography, ecology, and reproductive strategies of fishes.

Course Type: Graduate
Term: Spring
Day/Time: MW 1:00-2:15

Laboratory for Ichthyology [Grad] (E&EB 565)

Laboratory and field studies of fish diversity, form, function, behavior, and classification. The course primarily involves study of museum specimens and of living and fossil fishes.
 

Course Type: Graduate
Term: Spring

Biological Oceanography [Grad] (E&EB 575)

Exploration of a range of coastal and pelagic ecosystems. Relationships between biological systems and the physical processes that control the movements of water and productivity of marine systems. Anthropogenic impacts on oceans, such as the effects of fishing and climate change. Includes three Friday field trips.

Course Type: Graduate
Term: Spring
Day/Time: TTh 11.35-12.50

Evolutionary Functional Genomics, Cell Types and Homology [Grad] (E&EB 610)

Functional genomics has opened the opportunity to assess the activity state of all genes in the genomes in a largely scalable way. Many cell types, tissues and characters can readily be assessed across many species (if one has enough money), leading to a new field of evolutionary or comparative functional genomics. At the same time this new field of data analysis can be used to address many deep issues in organismic evolution, like the evolution of cell types, the homology among cell types etc.

In this seminar we will review the current state of published literature as it pertains to the evolutionary analysis of transcriptomes and epigenetic marks and their bearing on issues of cell and tissue evolution and homology.

Location: Yale Systems Biology Institute on Yale West Campus, Building ISTC (aka 850 West Campus Drive) room 101

Course Type: Graduate
Term: Fall
Day/Time: Tuesday, 3:30 pm- 4:30 pm ( or 5:00 pm)

Tropical Field Ecology (E&EB 617)

This course is designed to give students first-hand knowledge of tropical biology and the issues surrounding conservation of biodiversity in a developing nation, through a combination of seminar-style discussions and a mandatory field trip over spring break. The emphasis is on active learning and developing independent research projects carried out during the field trip. Using a case-study approach, topics covered include patterns of biodiversity, tropical forest dynamics, reforestation, species interactions and coevolution, climate change impacts, ecosystem services, and human land-use. Students also gain experience with study design, data collection methods, and statistical analysis. This year’s field trip is to Ecuador, a country famous for its high biological, cultural, and economic diversity. We visit a variety of forest ecosystems and hear from local and international scientists about current research in the field. Students undertake two short research projects and also learn basic identification and natural history of tropical plant, bird, and insect species. Students should expect to spend a major part of each day outside in the natural tropical environment under adverse conditions. Enrollment limited to fifteen. Simon A. Queenborough, Walter Jetz.

Course Type: Graduate
Term: Spring

Advanced Ecology [Grad] (E&EB 620)

An advanced treatment of ecology, including species interactions, species coexistence theory, species-environment interactions, the maintenance and consequences of biological diversity, spatial ecology, food webs, and eco-evolutionary interactions.

Course Type: Graduate
Term: Spring
Day/Time: T Th 1:00 - 2:15

Biology of Insect Disease Vectors [Grad] (E&EB 650)

Insects transmit pathogens that cause many emerging and re-emerging human and agriculture-related diseases. Many of these diseases, which are referred to as neglected tropical diseases (NTDs), have a dramatically negative impact on human health in the developing world. Furthermore, they cause indirect devastation by significantly reducing agricultural productivity and nutrient availability, exacerbating poverty and deepening disparities. This course introduces students to the biological interactions that occur between major groups of important disease vectors and the pathogens they transmit. Lectures cover current research trends that relate to the ecology and physiology of insect vectors. Course content focuses on how these aspects of vector biology relate to the development and implementation of innovative and effective disease-control strategies.

Prerequisite: full year of college/university-level biology, or permission of the instructor(s).

Course Type: Graduate
Term: Spring
Day/Time: W 3.00-4.20, Th 3.00-3.50

Ornithology [Grad] (E&EB 672)

An overview of avian biology and evolution, including the structure, function, behavior, and diversity of birds. The evolutionary origin of birds, avian phylogeny, anatomy, physiology, neurobiology, breeding systems, and biogeography.

Course Type: Graduate
Term: Spring
Day/Time: MWF 9.25-10.15

Laboratory for Ornithology [Grad] (E&EB 673)

Laboratory and field studies of avian morphology, diversity, phylogeny, classification, identification, and behavior.

Course Type: Graduate
Term: Spring
Day/Time: T 1.30-4.30

Life History Evolution [Grad] (E&EB 680)

Life history evolution studies how the phenotypic traits directly involved in reproductive success are shaped by evolution to solve ecological problems. Nowhere is the interplay between evolution and ecology more intimate.

Requires Permission of the Instructor

Course Type: Graduate
Term: Spring
Day/Time: TTh 11.35-12.50

Long-term Temporal Dynamics of Ecological Systems [Grad] (E&EB 740)

Are ecological systems generally at (or near) equilibrium? Or are their transient dynamics so slow that we need to know how they behave far from equilibrium, too? This question is increasingly pressing in the face of ongoing global change – there remains substantial uncertainty about whether predictions based on ecological equilibria are relevant for predicting ecosystem responses to global change. For insight into this question, we will deal with temporal dynamics of ecosystems, integrating theoretical perspectives with both modern long-term ecological research and paleoecology. We will consider how theoretical concepts like characteristic time scales, lag and legacy effects, and cyclic behaviors apply to empirical work in real ecological systems.

Course Type: Graduate
Term: Spring
Day/Time: Th 10:00-12:00

Evolving Dynamical Systems [Grad] (E&EB 810)

An introduction to the ways evolving biological systems can be described, modeled, and analyzed by using a dynamical systems approach.  Concrete models will be explored with respect to field or laboratory observations.  Extensive use will be made of the software package Mathematica, but prior experience with the program is not required.

Course Type: Graduate
Term: Fall
Day/Time: TTh 9.25-10.15

Primate Diversity & Evolution [Grad] (E&EB 842)

The diversity and evolutionary history of living and extinct primates.  Focus on major controversies in primate systematics and evolution, including the origins and relationships of several groups.  Consideration of both morphological and molecular studies.  Morphological and diversity and adaptations explored through museum specimens and fossil casts.

Course Type: Graduate
Term: Spring
Day/Time: W 1:30-3:20

1st Year Intro Research & Rotation [Grad] (E&EB 900)

1st Year Intro Research & Rotation

Course Type: Graduate
Term: Fall
Day/Time: 3 HTBA

2nd Year Research [Grad] (E&EB 950)

By arrangement with faculty.

Course Type: Graduate
Term: Spring , Term: Fall
Day/Time: 3 HTBA

Studies in Evolutionary Medicine I [Grad] (E&EB 960 )

The first term of a two-term course that begins in January. Students learn the major principles of evolutionary biology and apply them to issues in medical research and practice by presenting and discussing original papers from the current research literature. Such issues include lactose and alcohol tolerance; the hygiene hypothesis and autoimmune disease; human genetic variation in drug response and pathogen resistance; spontaneous abortions, immune genes, and mate choice; parental conflicts over reproductive investment mediated by genetic imprinting; life history trade-offs and the evolution of aging; the evolution of virulence and drug resistance in pathogens; the evolutionary genetics of humans and their pathogens; the ecology and evolution of disease; the evolutionary origin of diseases; and the emergence of new diseases. Students develop a research proposal based on one of their own questions in the spring term, spend the summer on a research project related to their research proposal, and write a paper based on the results of their research in the fall term. Credit and grades are awarded for each term. Only students who have engaged in summer research projects may enroll in the fall term.

Admission is by competitive application only.

http://eeb.yale.edu/sites/default/files/eeb_460b.pdf

Course Type: Graduate
Term: Spring
Day/Time: TTh 4.00-5.15

Studies in Evolutionary Medicine II [Grad] (E&EB 961)

Continuation of E&EB 960b. Students learn the major principles of evolutionary biology and apply them to issues in medical research and practice by presenting and discussing original papers from the current research literature. Such issues include lactose and alcohol tolerance; the hygiene hypothesis and autoimmune disease; human genetic variation in drug response and pathogen resistance; spontaneous abortions, immune genes, and mate choice; parental conflicts over reproductive investment mediated by genetic imprinting; life history trade-offs and the evolution of aging; the evolution of virulence and drug resistance in pathogens; the evolutionary genetics of humans and their pathogens; the ecology and evolution of disease; the evolutionary origin of diseases; and the emergence of new diseases. Students develop a research proposal based on one of their own questions in the spring term, spend the summer on a research project related to their research proposal, and write a paper based on the results of their research in the fall term. Credit and grades are awarded for each term. Only students who have engaged in summer research projects may enroll in the fall term.

Course Type: Graduate
Term: Fall
Day/Time: TTh 4.00-5.15